{"id":221,"date":"2018-07-17T10:43:22","date_gmt":"2018-07-17T14:43:22","guid":{"rendered":"http:\/\/site.caes.uga.edu\/carpophiline-id\/?page_id=221"},"modified":"2021-01-20T06:48:18","modified_gmt":"2021-01-20T11:48:18","slug":"urophorus-humeralis-fabricius-1798","status":"publish","type":"page","link":"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/","title":{"rendered":"Urophorus humeralis (Fabricius, 1798)"},"content":{"rendered":"\n<h3 class=\"wp-block-heading\"><strong>Species Fact Sheet<\/strong><br><em>Urophorus humeralis<\/em> (Fabricius, 1798)<\/h3>\n\n\n\n<div class=\"wp-block-image captionunderimg\"><figure class=\"alignright size-large is-resized\"><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal.jpg\"><img loading=\"lazy\" decoding=\"async\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal-1024x788.jpg\" alt=\"\" class=\"wp-image-467\" width=\"300\" height=\"231\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal-1024x788.jpg 1024w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal-300x231.jpg 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal-768x591.jpg 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal-179x138.jpg 179w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/HumeralisMaleHabitusFinal.jpg 1200w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/><\/a><figcaption><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 1. Male dorsal and ventral habitus<\/p><\/figcaption><\/figure><\/div>\n\n\n<p style=\"text-align: left\"><strong>Diagnosis:<\/strong><em> Urophorus humeralis<\/em> specimens are distinguished from other eastern North American species by having three tergites visible beyond the elytra in dorsal view (Fig. 1). Males also have a round punctate depression on the 5th ventrite anterior to the supplementary segment and distinctly longer setae located medially on the posterior margin of the 4th ventrite (Fig. 1 &amp; 8). Females are distinguished by a large round depression on the pygidium (Fig. 7).&nbsp;<\/p>\n<p style=\"text-align: left\"><strong>Note:<\/strong> The number of visible tergites in Carpophilinae specimens may be misleading in some specimens due to distention or retraction caused by certain preservation methods (Fig. 9). This occurs commonly in <em>Urophorus humeralis<\/em> specimens. The specimen in Fig. 9 has the 2nd tergite distended and the 5th tergite (pygidium) retracted. Distended tergites can be distinguished from those which are normally visible by only being sclerotized in the posterior half only.<\/p>\n<p style=\"text-align: left\"><strong>Distribution:<\/strong> <em>Urophorus humeralis<\/em> is a cosmopolitan species and one of the most widely spread Carpophilinae species. Thought to be native to Asia, it is now spread to every continent. Found throughout the US.<\/p>\n<p style=\"text-align: left\"><strong>Biology:<\/strong> <em>U. humeralis<\/em> can be found feeding on most fruits and vegetables, especially when they are over ripened. This species is also known to infest stored food products and can be a pest in processing plants (Schmidt, 1935). <em>U. humeralis<\/em> is known to transport pathogens from one fruit to another. Develops from egg to adult in 29 days at 24\u2103 (Connell, 1956). Larvae illustrated and described by Connell, 1956.<\/p>\n<p style=\"text-align: left\"><strong>References:<\/strong><br>(1) Chang, V. C. S., &amp; Jensen, L. (1974) \u2013 Transmission of pathogens in pineapples<br>(2) Connell, W. A. (1956) \u2013 larvae and adult descriptions and key to larvae and adults<br>(3) Denux O., Zagatti P. (2010) \u2013 Distribution<br>(4) Schmidt, C. T. (1935) \u2013 feeding habits and development rates<\/p>\n\n\n<p class=\"has-text-align-center has-small-font-size\"><strong>Double-click on images to enlarge<\/strong><\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter columns-3 is-cropped captionunderimg wp-block-gallery-1 is-layout-flex wp-block-gallery-is-layout-flex\"><ul class=\"blocks-gallery-grid\"><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae.png\"><img loading=\"lazy\" decoding=\"async\" width=\"835\" height=\"557\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae.png\" alt=\"\" data-id=\"516\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/uhumeralisantennae-3\/\" class=\"wp-image-516\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae.png 835w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae-300x200.png 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae-768x512.png 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisAntennae-207x138.png 207w\" sizes=\"auto, (max-width: 835px) 100vw, 835px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\">Figure <p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">2. Antenna (dorsal view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum.png\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"788\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum-1024x788.png\" alt=\"\" data-id=\"514\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/humeralismalepronotum-2\/\" class=\"wp-image-514\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum-1024x788.png 1024w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum-300x231.png 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum-768x591.png 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum-179x138.png 179w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMalePronotum.png 1145w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 3. Pronotum (dorsal view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge.png\"><img loading=\"lazy\" decoding=\"async\" width=\"568\" height=\"829\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge.png\" alt=\"\" data-id=\"517\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/uhumeralismesosternumridge-3\/\" class=\"wp-image-517\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge.png 568w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge-206x300.png 206w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/Uhumeralismesosternumridge-95x138.png 95w\" sizes=\"auto, (max-width: 568px) 100vw, 568px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 4. Mesosternum with anterior impunctate edge along with median longitudinal ridge (ventral view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1.png\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1-1024x683.png\" alt=\"\" data-id=\"426\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/uhumeralisaxillaryspace-2\/\" class=\"wp-image-426\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1-1024x683.png 1024w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1-300x200.png 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1-768x512.png 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1-207x138.png 207w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2018\/07\/UhumeralisAxillaryspace-1.png 1074w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 5. Posterior rim of mesocoxal cavities (ventrolateral view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet.jpg\"><img loading=\"lazy\" decoding=\"async\" width=\"371\" height=\"542\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet.jpg\" alt=\"\" data-id=\"515\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet.jpg\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/maletibiafactsheet\/\" class=\"wp-image-515\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet.jpg 371w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet-205x300.jpg 205w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/maletibiafactsheet-94x138.jpg 94w\" sizes=\"auto, (max-width: 371px) 100vw, 371px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\">Figure <p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">6. Male metathoracic tibia shape, gradually dilated (anterior view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex.png\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex-1024x683.png\" alt=\"\" data-id=\"512\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/humeralisfemalabdomenapex-2\/\" class=\"wp-image-512\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex-1024x683.png 1024w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex-300x200.png 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex-768x512.png 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex-207x138.png 207w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisFemalAbdomenApex.png 1200w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 7. Female pygidium (dorsal view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep.png\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"683\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep-1024x683.png\" alt=\"\" data-id=\"513\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/humeralismaledep-2\/\" class=\"wp-image-513\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep-1024x683.png 1024w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep-300x200.png 300w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep-768x512.png 768w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep-207x138.png 207w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/HumeralisMaleDep.png 1200w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 8. Male supplementary segment (ventral view)<\/p><\/figcaption><\/figure><\/li><li class=\"blocks-gallery-item\"><figure><a href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical.png\"><img loading=\"lazy\" decoding=\"async\" width=\"683\" height=\"1024\" src=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical-683x1024.png\" alt=\"\" data-id=\"521\" data-full-url=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical.png\" data-link=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/urophorus-humeralis-fabricius-1798\/uhumeralishabitusmaledorsalvertical-4\/\" class=\"wp-image-521\" srcset=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical.png 683w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical-200x300.png 200w, https:\/\/site.caes.uga.edu\/carpophiline-id\/files\/2019\/07\/UhumeralisHabitusmaledorsalvertical-92x138.png 92w\" sizes=\"auto, (max-width: 683px) 100vw, 683px\" \/><\/a><figcaption class=\"blocks-gallery-item__caption\"><p style=\"color: #ffffff;background: #000000;border: 4px solid #dddddd\">Figure 9. Male with 2nd tergite distended and the 5th tergite (pygidium) retracted (dorsal view)<\/p><\/figcaption><\/figure><\/li><\/ul><\/figure>\n\n\n\n<p><strong>HOW TO CITE THIS WORK:<\/strong><\/p>\n\n\n\n<p>DiLorenzo, C.L., G.S. Powell, A.R. Cline, and J.V. McHugh (2021) Carpophiline-ID, a taxonomic web resource for the identification of Carpophilinae (Nitidulidae) of eastern North America. (vers. 01.19.2021) University of Georgia, retrieved from\u00a0<a rel=\"noreferrer noopener\" href=\"https:\/\/site.caes.uga.edu\/carpophiline-id\/\" target=\"_blank\">https:\/\/site.caes.uga.edu\/carpophiline-id\/<\/a><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Species Fact SheetUrophorus humeralis (Fabricius, 1798) Diagnosis: Urophorus humeralis specimens are distinguished from other eastern North American species by having three tergites visible beyond the elytra in dorsal view (Fig. 1). Males also have a round punctate depression on the 5th ventrite anterior to the supplementary segment and distinctly longer setae located medially on the [&hellip;]<\/p>\n","protected":false},"author":147,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"page-full.php","meta":{"footnotes":""},"class_list":["post-221","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/pages\/221","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/users\/147"}],"replies":[{"embeddable":true,"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/comments?post=221"}],"version-history":[{"count":9,"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/pages\/221\/revisions"}],"predecessor-version":[{"id":1493,"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/pages\/221\/revisions\/1493"}],"wp:attachment":[{"href":"https:\/\/site.caes.uga.edu\/carpophiline-id\/wp-json\/wp\/v2\/media?parent=221"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}